StegocephaliStegocephali (often spelled Stegocephalia) is a group containing all four-limbed vertebrates. It is equivalent to a broad definition of Tetrapoda: under this broad definition, the term "tetrapod" applies to any animal descended from the first vertebrate with limbs and toes, rather than fins. This includes both the modern lineage of limbed vertebrates (the crown group, including modern amphibians, mammals, reptiles and birds) as well as a portion of the stem group, limbed vertebrates that evolved prior to the origin of the crown group.
Carboniferous rainforest collapseThe Carboniferous rainforest collapse (CRC) was a minor extinction event that occurred around 305 million years ago in the Carboniferous period. It altered the vast coal forests that covered the equatorial region of Euramerica (Europe and America). This event may have fragmented the forests into isolated refugia or ecological 'islands', which in turn encouraged dwarfism and, shortly after, extinction of many plant and animal species. Following the event, coal-forming tropical forests continued in large areas of the Earth, but their extent and composition were changed.
Romer's gapRomer's gap is an example of an apparent gap in the tetrapod fossil record used in the study of evolutionary biology. Such gaps represent periods from which excavators have not yet found relevant fossils. Romer's gap is named after paleontologist Alfred Romer, who first recognised it. Recent discoveries in Scotland are beginning to close this gap in palaeontological knowledge. Romer's gap ran from approximately 360 to 345 million years ago, corresponding to the first 15 million years of the Carboniferous, the early Mississippian (starting with the Tournaisian and moving into the Visean).
Evolutionary gradeA grade is a taxon united by a level of morphological or physiological complexity. The term was coined by British biologist Julian Huxley, to contrast with clade, a strictly phylogenetic unit. An evolutionary grade is a group of species united by morphological or physiological traits, that has given rise to another group that has major differences from the ancestral condition, and is thus not considered part of the ancestral group, while still having enough similarities that we can group them under the same clade.
AnamniotesThe anamniotes are an informal group of craniates comprising all fishes and amphibians, which lay their eggs in aquatic environments. They are distinguished from the amniotes (reptiles, birds and mammals), which can reproduce on dry land either by laying shelled eggs or by carrying fertilized eggs within the female. Older sources, particularly before the 20th century, may refer to anamniotes as "lower vertebrates" and amniotes as "higher vertebrates", based on the antiquated idea of the evolutionary great chain of being.
Skull roofThe skull roof, or the roofing bones of the skull, are a set of bones covering the brain, eyes and nostrils in bony fishes and all land-living vertebrates. The bones are derived from dermal bone and are part of the dermatocranium. In comparative anatomy the term is used on the full dermatocranium. In general anatomy, the roofing bones may refer specifically to the bones that form above and alongside the brain and neurocranium (i.e.
Vertebral columnThe vertebral column, also known as the backbone or spine, is part of the axial skeleton. The vertebral column is the defining characteristic of a vertebrate in which the notochord (a flexible rod of uniform composition) found in all chordates has been replaced by a segmented series of bone: vertebrae separated by intervertebral discs. Individual vertebrae are named according to their region and position, and can be used as anatomical landmarks in order to guide procedures such as lumbar punctures.
Permian–Triassic extinction eventThe Permian–Triassic (P–T, P–Tr) extinction event (PTME), also known as the Late Permian extinction event, the Latest Permian extinction event, the End-Permian extinction event, and colloquially as the Great Dying, forms the boundary between the Permian and Triassic geologic periods, and with them the Paleozoic and Mesozoic eras respectively, approximately 251.9 million years ago. As the largest of the "Big Five" mass extinctions of the Phanerozoic, it is the Earth's most severe known extinction event, with the extinction of 57% of biological families, 83% of genera, 81% of marine species and 70% of terrestrial vertebrate species.
VertebraThe spinal column, a defining synapomorphy shared by nearly all vertebrates, is a moderately flexible series of vertebrae (: vertebra), each constituting a characteristic irregular bone whose complex structure is composed primarily of bone, and secondarily of hyaline cartilage. They show variation in the proportion contributed by these two tissue types; such variations correlate on one hand with the cerebral/caudal rank (i.e., location within the backbone), and on the other with phylogenetic differences among the vertebrate taxa.
AmnioteAmniotes belong to the clade Amniota, a clade of tetrapod vertebrates that comprises sauropsids (including all reptiles and birds) and synapsids (including mammals and mammal ancestors like "pelycosaurs" and therapsids). They are distinguished from the other living tetrapod clade—the lissamphibians—by the development of three extraembryonic membranes (amnion for embryoic protection, chorion for gas exchange, and allantois for metabolic waste disposal or storage), thicker and more keratinized skin, and costal respiration (breathing by expanding/constricting the rib cage).
EndocraniumThe endocranium in comparative anatomy is a part of the skull base in vertebrates and it represents the basal, inner part of the cranium. The term is also applied to the outer layer of the dura mater in human anatomy. Structurally, the endocranium consists of a boxlike shape, open at the top. The posterior margin exhibit the foramen magnum, an opening for the spinal cord. The floor of the endocranium has several paired openings for the cranial nerves, and the anterior margin holds a spongy construction, allowing for the external nasal nerves to pass through.
Transitional fossilA transitional fossil is any fossilized remains of a life form that exhibits traits common to both an ancestral group and its derived descendant group. This is especially important where the descendant group is sharply differentiated by gross anatomy and mode of living from the ancestral group. These fossils serve as a reminder that taxonomic divisions are human constructs that have been imposed in hindsight on a continuum of variation.
Evolution of fishThe evolution of fish began about 530 million years ago during the Cambrian explosion. It was during this time that the early chordates developed the skull and the vertebral column, leading to the first craniates and vertebrates. The first fish lineages belong to the Agnatha, or jawless fish. Early examples include Haikouichthys. During the late Cambrian, eel-like jawless fish called the conodonts, and small mostly armoured fish known as ostracoderms, first appeared.
SynapsidSynapsids are one of the two major clades of vertebrate animals that evolved from basal amniotes, the other being the sauropsids, which include reptiles (turtles, crocodilians and lepidosaurs) and birds. The synapsids were once the dominant land animals in the late Paleozoic and early Mesozoic, but the only extant group that survived into the Cenozoic are the mammals. Unlike other amniotes, synapsids have a single temporal fenestra, an opening low in the skull roof behind each eye orbit, leaving a bony arch beneath each; this accounts for their name.
Vertebrate paleontologyVertebrate paleontology is the subfield of paleontology that seeks to discover, through the study of fossilized remains, the behavior, reproduction and appearance of extinct animals with vertebrae or a notochord. It also tries to connect, by using the evolutionary timeline, the animals of the past and their modern-day relatives. The fossil record shows aspects of the meandering evolutionary path from early aquatic vertebrates to mammals, with a host of transitional fossils, though there are still large blank areas.
EusthenopteronEusthenopteron (from εὖ eû, 'good', σθένος sthénos, 'strength', and πτερόν pteron 'wing' or 'fin') is a genus of prehistoric sarcopterygian (often called lobe-finned fishes) which has attained an iconic status from its close relationships to tetrapods. Early depictions of this animal show it emerging onto land; however, paleontologists now widely agree that it was a strictly aquatic animal. The genus Eusthenopteron is known from several species that lived during the Late Devonian period, about 385 million years ago.
FishA fish (: fish or fishes) is an aquatic, craniate, gill-bearing animal that lacks limbs with digits. Included in this definition are the living hagfish, lampreys, and cartilaginous and bony fish as well as various extinct related groups. Approximately 95% of living fish species are ray-finned fish, belonging to the class Actinopterygii, with around 99% of those being teleosts. The earliest organisms that can be classified as fish were soft-bodied chordates that first appeared during the Cambrian period.
LungfishLungfish are freshwater vertebrates belonging to the class Dipnoi. Lungfish are best known for retaining ancestral characteristics within the Osteichthyes, including the ability to breathe air, and ancestral structures within Sarcopterygii, including the presence of lobed fins with a well-developed internal skeleton. Lungfish represent the closest living relatives of the tetrapods. The mouths of lungfish typically bear tooth plates, which are used to crush hard shelled organisms.
